The Medrash that I quoted earlier actually says ???? ????? ????? ??? ??? ????? ??? ????? ????? ??? ????? ???? ?? ???? ????? ?? ???? ???? ????? ??? ????? ???? which lichora means that they shevatim married their full sisters. The Mateh Efraim in fact asks this as a kashya on those who explained that the shevatim married each other’s twins.
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